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Upper Ordovician carbon isotope chemostratigraphy and a high-resolution assessment of the Hirnantian Stage in the Baltic Sea subsurface

ABSTRACT Based on a new deep drilling on southern Gotland (Sweden), this study is the first to document the carbon isotope chemostratigraphy of the Upper Ordovician through lowermost Silurian sedimentary record from the central parts of the Baltic Sea subsurface. The lithological record of the Stora Sutarve core has similarities with adjacent successions in both the Viki core in the East Baltic area and the Borenshult core in the Swedish mainland. The core includes the Kahula (including 13 bentonites in the “Kinnekulle K-bentonite complex”), Hirmuse(?), Rägavere, Paekna, Jonstorp, Loka, Motala and Kallholn formations. Several of the internationally recognized upper Ordovician δ13C excursions have been identified in the core, including the Guttenberg Isotope Carbon Excursion (GICE), Moe excursion, Hirnantian Isotope Carbon Excursion (HICE), and an undefined early Silurian carbon isotope excursion, presumably the Early Aeronian Carbon Isotope Excursion (EACIE). We particularly address the microstratigraphy of the Hirnantian Stage and the finer details of the HICE, which is not fully complete and entirely confined to the 2.09-m-thick Loka Formation. This formation yields abundant, but low-richness brachiopod faunas characteristic of the Hirnantian Stage and is, based on erosional-depositional surfaces and facies, interpreted as reflecting interglacial warming and eustatic transgression related to ice-sheet contraction in Gondwana. Based on very dense carbon isotope sampling we identify four clusters of δ13C values that aid to define significant changes in the preserved record of the HICE and which may facilitate global correlation of the succession and the herein interpreted sea-level record.

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Thin loess in Southwestern Sweden

ABSTRACT A thin (20–80 cm), patchy layer of silt-rich sediment occurs at the surface throughout Svartedalen, a nature reserve 30 km north of Gothenburg, Sweden. This surface silt mantles a bedrock-dominated, fracture-valley landscape. Using data from grain-size analysis, OSL dating and detrital-zircon U-Pb dating, we argue that the silt is loess sourced from glacial sediment that was eroded from local bedrock. The sediment has a grain-size distribution typical of wind-blown silt and is especially similar to thin deposits of loess overlying coarser material. OSL ages on five samples range from 1 to 8 ka, although analysis of equivalent dose distributions of one may suggest an age as old as 11 ka. The dates may represent true depositional ages and represent several Holocene eolian events. However, we consider as more likely that the loess was deposited during deglaciation, and quartz-grain signals have been partially reset during bioturbation. U-Pb ages on 273 zircon grains from the loess show prominent peaks at 1.6 and 1.8 Ga, as well as smaller numbers of grains from 1.0 to 1.6 Ga. These ages match dates from the Idefjord Terrane which comprises the bedrock of the study area. We argue that during ice-margin retreat, debris in the glacier was dominated by locally derived debris. This glacial sediment was left in thin patches uplands and particularly in large ice-marginal deltas. These deposits served as the proximal source for the loess. The presence of thin loess in Svartedalen suggests loess to be common in soils of southwest Sweden.

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Shell injuries, repair and malformation in the early Cambrian mollusc Helcionella antiqua from Scania, Sweden

ABSTRACT Three cases of repaired injuries and malformation in specimens of the helcionelloid mollusc Helcionella antiqua (Kiær, 1917) from the lower Cambrian (Cambrian Series 2, Stage 4) Gislöv Formation of southern Sweden document some of the oldest known durophagous attacks on Palaeozoic molluscs. Two of the injuries are developed as clefts, of which one had a severe effect on the continued growth of the shell. The third example is a large embayment removing large portions of the supra-apical part of the shell. A similar repaired injury is known in the slightly older mollusc Marocella mira Geyer, 1986. from Antarctica and Australia. The morphology of the injuries and the hydrodynamically quiet depositional setting suggests that the shell damage was caused by failed predatory attacks. The location of the repaired injuries suggests that the attacks may have targeted the head region of the molluscs, thus supporting an endogastrically coiled orientation of the shell in Helcionella. Only three repaired injuries in 252 Helcionella specimens were found, giving a shell repair frequency of 1.2%. All three examples occur in the larger size classes. The size-frequency distribution (N = 182) is strongly right skewed, which could suggest high input of juvenile specimens into the assemblage. The assemblage is interpreted as a time averaged and mixed death assemblage, albeit with good correspondence with the living shelly assemblage, due to a relatively thin, homogenous unit that may suggest within-habitat time averaging.

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Pore-canal network (“wrinkles”) in ammonoid shell wall (Cephalopoda)

ABSTRACT The shell wall in ammonoids contains a pore-canal network similar to that detected recently in the shell wall of the Ordovician nautiloid Orthoceras. In the Jurassic monomorphic ammonoids Quenstedtoceras, Cosmoceras, Lissoceras and the Cretaceous heteromorphic ammonoid Ptychoceras investigated in this study vertical canals arranged in parallel rows, are partially preserved, although no horizontal canals are preserved as in Orthoceras. In Quenstedtoceras and Ptychoceras the vertical canals extend not only through the inner prismatic layer, as in Orthoceras, but also through the nacreous layer. Therefore both layers were secreted on the body-shell attachment band behind the apertural margin. In the juvenile shells, only the very thin outer prismatic layer formed the apertural margin. As in Orthoceras, the canals had walls of calcified organic fibres that were partially or entirely dissolved during diagenesis. The pore-canal network in the shell wall of Quenstedtoceras and Ptychoceras contributed to achieving neutral buoyancy. As indicated by the high numbers of mini-pores in the connecting rings, the siphuncle was employed for adjusting buoyancy during vertical migrations. The “wrinkles”, “furrows”, “pits”, “dots”, “ridges”, “Ritzsteifen” and “Runselschicht” in the “wrinkle layer”, previously described in the ammonoid shell walls, are remnants of the pore-canal network.

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High diversity and early radiation of organic-walled phytoplankton in southern Baltica during the Middle-Late Ordovician – evidence from the Borenshult-1 drillcore of southern Sweden

ABSTRACT Highly diverse and well preserved organic-walled phytoplankton were recorded from the Darriwilian–early Katian interval of the Borenshult-1 drillcore. We identified 154 species in 53 genera, and three assemblages were distinguished; Assemblage A of a late Darriwilian age, Assemblage B of a Sandbian age (further subdivided into sub-assemblages B1 and B2), and Assemblage C dated as Katian. Taxa with “Silurian affinities” with previous first appearance datum in the early Silurian, Hirnantian, such as Metaleiofusa and Visbysphaera, are here recorded from the late Darriwilian and Sandbian respectively. These occurrences question the relationship between the appearance of pioneering phytoplankton morphotypes and the Hirnantian glaciation. Other taxa with no pre-Silurian records such as Visbysphaera pirifera subsp. minor, Petaloferidium cazurrum and Dorsennidium cf. D. estrellitae are here present in the Sandbian, where bentonite beds are intercalated. The diversity curve of acritarchs shows similarities with those proposed for the Darriwilian-Katian of Baltica with main differences in the interval with bentonite beds representing an intense volcanic activity. The species Metaleiofusa arcuata Wall is here emended and a new combination is proposed: Petaloferidium cazurrum (Cramer) comb. nov. The genus Fankea is recorded for the first time from Swedish strata, suggesting a dominant high- to middle latitudinal distribution instead of a Perigondwanan distribution. We contend that the location of paleo-southern Sweden contributed to the great diversity seen, since a middle-low latitude provided a suitable habitat with warm, shallow water, rich in nutrients.

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